| Main Ref. | Yoneda, M., M. Tokimura, H. Fujita, N. Takeshita, K. Takeshita, M. Matsuyama and S. Matsuura, 2001 |
| Mode | dioecism |
| Fertilization | external |
| Mating type | |
| Spawning frequency | one clear seasonal peak per year |
| Batch spawner | Yes. Ref. Yoneda, M., M. Tokimura, H. Fujita, N. Takeshita, K. Takeshita, M. Matsuyama and S. Matsuura, 2001 |
| Reproductive guild |
nonguarders open water/substratum egg scatterers |
| Parental Care | none |
| Description of life cycle and mating behavior | |
| Search for more references on reproduction | Scirus |