Actinopterygii (ray-finned fishes) >
Perciformes (Perch-likes) >
Pholidae (Gunnels)
Etymology: Pholis: Greek, pholis, -idos = scale (Ref. 45335).
Environment / Climate / Range
Ecology
Marine; brackish; demersal; non-migratory; depth range 0 - 100 m (Ref. 48978), usually 0 - 30 m (Ref. 35388). Polar, preferred ?; 72°N - 43°N, 76°W - 45°E
North Atlantic: Kanin Peninsula southward to La Rochelle (including White Sea, North Sea and Baltic), also Iceland, but not eastern Greenland, Spitzbergen or Novaya Zemlya. Western Atlantic: Labrador, Canada to Delaware Bay, USA (Ref. 7251).
Size / Weight / Age
Maturity: Lm ?  range ? - ? cm
Max length : 25.0 cm SL male/unsexed; (Ref. 4693); max. reported age: 5 years (Ref. 72462)
Short description
Morphology | Morphometrics
Ribbon - shaped body with 9 - 13 eye spots along the base of the dorsal fin (Ref. 35388).
Occurs in shallow waters (seashore, tide pools), but descends (especially in winter) to 100 m or more. May remain out of water under rocks or seaweeds (Ref. 31184). Feeds on small crustaceans, polychaetes, mollusks and fish eggs. The eggs are laid on the sea bed in a large clump, which is closely guarded until hatched (Ref. 9900). Breathes air when out of water (Ref. 31184). Spawning occur in November - January . Female lays 80-200 eggs in a large ball under a stone or in an empty bivalve shell (Ref. 35388).
Male and female coil around the egg ball (Ref. 58332).
Makushok, V.M., 1986. Pholididae. p. 1124-1125. In P.J.P. Whitehead, M.-L. Bauchot, J.-C. Hureau, J. Nielsen and E. Tortonese (eds.) Fishes of the North-eastern Atlantic and the Mediterranean. UNESCO, Paris. Vol. 3. (Ref. 4693)
IUCN Red List Status (Ref. 115185)
CITES (Ref. 94142)
Not Evaluated
Threat to humans
Harmless
Human uses
Fisheries: of no interest; aquarium: commercial
More information
ReferencesAquacultureAquaculture profileStrainsGeneticsAllele frequenciesHeritabilityDiseasesProcessingMass conversion
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Estimates of some properties based on models
Phylogenetic diversity index (Ref.
82805): PD
50 = 0.5005 [Uniqueness, from 0.5 = low to 2.0 = high].
Bayesian length-weight: a=0.00219 (0.00096 - 0.00497), b=3.14 (2.94 - 3.34), in cm Total Length, based on LWR estimates for this species & (Sub)family-body (Ref.
93245).
Trophic Level (Ref.
69278): 3.5 ±0.46 se; Based on food items.
Resilience (Ref.
69278): Medium, minimum population doubling time 1.4 - 4.4 years (tm=1-3; K=0.18-0.29; tmax=12).
Prior r = 1.11, 2 SD range = 0.66 - 1.85, log(r) = 0.1, SD log(r) = 0.26, Based on: 3 K, 2 tgen, 1 tmax, records
Vulnerability (Ref.
59153): Low to moderate vulnerability (34 of 100) .