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| Sound produced | yes, active sound production |
| Type of sound produced | knocks, single or pulsed drum beats |
| Sound production organ | swim bladder |
| Sound mechanism | vibration by contraction of primary sonic muscles & conformation of swim bladder |
| Behavioural context | intraspecific competition, terretorial behaviour |
| Reference | Parmentier, E., P. Vandewalle and J.P. Lagardère, 2003 |
| Remark | strongly species-specific sounds, aquarium recording, recorded with lowpass filter: 6400 Hz. |