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| Sound produced | yes, active sound production |
| Type of sound produced | grunts |
| Sound production organ | swim bladder & pharyngeal teeth |
| Sound mechanism | vibration of swim bladder by general body contraction and striking of opercula against body, stridulatory sounds probably generated by teeth & amplified by adjacent swim bladder |
| Behavioural context | under duress (mild manual or electrical stimulation) |
| Reference | Fish, M.P. and W.H. Mowbray, 1970 |
| Remark | recording amplified by 6 dB |