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| Sound produced | yes, active sound production |
| Type of sound produced | thumps & knocks |
| Sound production organ | pharyngeal teeth & swim bladder |
| Sound mechanism | pharyngeal teeth stridulation augmented by adjacent swim bladder |
| Behavioural context | spontaneous sounds during competitive feeding |
| Reference | Fish, M.P. and W.H. Mowbray, 1970 |
| Remark | recording made in outdoor pen with bandpass filter: 30 - 1200 Hz, high pass noise reduction filter applied to recording & amplified by 3 dB |