Family Ceratiidae - Warty Seadevils
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Order |
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Class |
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Actinopterygii |
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No. in FishBase |
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Genera : 2 | Species : 4 Eschmeyer's Catalog of Fishes |
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Environment |
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Fresh : No |
Brackish : No |
Marine : Yes |
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Division |
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Marine |
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Aquarium |
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none |
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First Fossil Record |
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Remark |
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From Pietsch (2009). Distribution: Atlantic, Indian and Pacific Oceans. Distinguishing Character: Elongate, laterally compressed and relatively large in size with the cleft of the mouth vertical to strongly oblique. The back end of the peculiarly elongate pterygiophore of the illicium, emanating from the top midline of the trunk, retracts back and forth through a narrow cranial trough found along the full length of the dorsal region of the cranium, the posterior end of which extends sunken within a cavernous cylindrical skin evagination, projecting anterior to the caruncles. Metamorphosed females lacking sphenotic, quadrate, articular, angular, and preopercular spines. Illicium between the frontal bones on the snout. Skin completely covered with close-set dermal spinules. Notable unique feature are 2 - 3 caruncles (fleshy, wartlike glands) present before soft dorsal fin in females as a modification of 2 or 3 fin rays; soft rays usually 4 (rarely 5) in dorsal and anal fins.
Females are host to the sexually parasitic mature males. Larvae "hump-backed", the mouth somewhat vertical and skin moderately inflated. Well-developed sexual dimorphism in all stages, with distinct illicial rudiment and caruncles on the dorsal midline of the trunk. Small pectoral fins. Second dorsal spine of female larvae and juvenile having a distal bioluminescent gland that is subsequently reduced and hidden subcutaneously just behind the base of the illicium in adults. With 4 pectoral radials. Esca oval-shaped and may or may not have 1 - 2 distal appendages. Free-living metamorphosed males having notably large, bowl-shaped eyes and in contrast, having minute olfactory organs. At the tip of the snout is a pair of large denticular teeth, fused at the base and articulating with the pterygiophore of the illicium. At the tip of the lower jaw are 2 pairs of denticular teeth. In juvenile free-living stages, the skin is naked and devoid of pigmentation becoming spinulose and darkly pigmented in parasitic stages.
Ceratias holboelli females having standard length of at least 85.5 cm. Free-living males standard length of less than 0.2 cm. Parasitic males length range 0.08 - 14.0 cm. (Ref. 86949)
Comments: Dorsal-fin rays rarely 3 or 5, excluding the 2 or 3 reduced rays within the caruncles; pectoral-fin rays 15 - 19 (rarely 14); caudal-fin rays 8 or 9 (8 in Cryptopsaras; the ninth, lowermost ray reduced in Ceratias; 2 simple + 4 bifurcated + 2 (or 3) simple). (Ref. 86949) |
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Etymology |
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Greek, keras or keratos = horn or horned (in reference to the lure projecting from the snout) (Ref. 86949) |
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Reproductive guild |
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nonguarders |
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Typical activity level |
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Coordinator |
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Osteichthyes | Actinopterygii | Actinopteri | Neopterygii | Teleostei | Osteoglossocephalai | Clupeocephala | Euteleosteomorpha | Neoteleostei | Eurypterygia | Ctenosquamata | Acanthomorphata | Euacanthomorphacea | Percomorphaceae | Percomorpharia | Lophiiformes | Ceratioidei | Ceratiidae |
Important recommendation:
The list below must not be used as an authority reference synonymy list like those found in scientific published revisions, which must be the source to be used and cited eventually when they exist.
Rather, it reflects the current content of FishBase, and the progress with respect to synchronization with the Catalog of Fishes. However, we think it can be useful for users to assess the quality of information in FishBase, to start new work on the family, or to cross-check with other lists.
But we appreciate to be cited in publications when this list has been of any working value. In particular, for published scientific, we suggest then to cite it in the Material and Method section as a useful tool to conduct the research, but again, not as a taxonomic or nomenclatural authority reference.
Unless it is explicitly precised, the list is not complete, please search all original names published for the family in the Catalog of Fishes (genera, species), including those with uncertain or unknown status, that are not included in FishBase when they are not attached to a valid species.
This list uses some data from Catalog of Fishes (not shown but used to sort names).
In the column CofF, the digit indicates the status of synchronization with CofF: 0: Not checked; 1: Same status; 2: Different status; 3: Other combination; 4: Synonym in CofF; 5: Species/Subspecies issue; 6: Synonym of another species in CofF; 7: Not in CofF; 8: Should not be in CofF. The CofF version currently used is the one published on 23-07-2014 (Ref. 97102).
The list ordered as follows:
Please send comments and corrections if you detect errors or missing names.
Scientifc name | Status | Senior/Junior synonym | Combination | CofF |
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Ceratias holboelli Krøyer, 1845 | accepted | senior | original | 2 |
Reganula giganteus (Bigelow & Barbour, 1944) | synonym | junior | new | |
Ceratias uranoscopus Murray, 1877 | accepted | senior | original | 2 |
Manchalias uranoscopus (Murray, 1877) | synonym | senior | new | |
! Mancalias uranoscopus (Murray, 1877) | synonym | senior | new | |
Typlopsaras shufeldti Gill, 1883 | synonym | junior | original | 2 |
! Typhlopsaras shufeldti Gill, 1883 | synonym | junior | original | |
Mancalias xenistius Regan & Trewavas, 1932 | synonym | junior | original | 2 |
! Manchalias xenistius Regan & Trewavas, 1932 | synonym | junior | original | |
Ceratias holboelli (non Krøyer, 1845) | misapplied | misapplied | original | |
Ceratias tentaculatus (Norman, 1930) | accepted | senior | new | |
Mancalias tentaculatus Norman, 1930 | synonym | senior | original | 2 |
Ceratias holboelli tentaculatus (Norman, 1930) | synonym | senior | new | |
Mancalias bifilis Regan & Trewavas, 1932 | synonym | junior | original | 2 |
Ceratias holboelli (non Krøyer, 1845) | misapplied | misapplied | original | |
Cryptopsaras couesii Gill, 1883 | accepted | senior | original | 2 |
Ceratias couesii (Gill, 1883) | synonym | senior | new | |
! Cryptopsaras couesi Gill, 1883 | synonym | senior | original | |
! Cryptosaras couesi Gill, 1883 | synonym | senior | original | |
! Cryptosarus couesi Gill, 1883 | synonym | senior | original | |
Ceratias carunculatus Günther, 1887 | synonym | junior | original | 2 |
Cryptopsaras carunculatus (Günther, 1887) | synonym | junior | new | |
! Cryptosparas carunculatus (Günther, 1887) | synonym | junior | new | |
Ceratias mitsukurii Tanaka, 1908 | synonym | junior | original | 2 |
! Cryptosparas pennifer Regan & Trewavas, 1932 | synonym | junior | original | 2 |
Cryptopsaras pennifer Regan & Trewavas, 1932 | synonym | junior | original | |
! Cryptosparas valdiviae Regan & Trewavas, 1932 | synonym | junior | original | 2 |
Cryptopsaras valdiviae Regan & Trewavas, 1932 | synonym | junior | original | |
! Cryptosparas normani Regan & Trewavas, 1932 | synonym | junior | original | 2 |
Cryptopsaras normani Regan & Trewavas, 1932 | synonym | junior | original | |
! Cryptosparas atlantidis Barbour, 1941 | synonym | junior | original | 2 |
Cryptopsaras atlantidis Barbour, 1941 | synonym | junior | original |